Penstemon strictus Benth. ssp. angustus Pennell

Scientific Name: Penstemon strictus Benth. ssp. angustus Pennell


Classification: Plantae/ Tracheobionta / Spermatophyta / Magnoliophyta / Magnoliopsida / Asteridae / Scrophulariales / Scrophulariaceae / Penstemon Schmidel/ Penstemon strictus Benth. ssp. angustus Pennell

<i>Penstemon strictus</i> Benth. ssp. angustus Pennell
General Information
Usda SymbolPESTA
GroupDicot
Life CyclePerennial
Growth HabitsForb/herb
Native LocationsPESTA

Plant Guide

Alternate Names

Strict beardtongue, Rocky Mountain beardtongue

Uses

Grazing/rangeland: 'Cedar' Palmer’s penstemon (Penstemon palmeri) and 'Bandera' Rocky Mountain penstemon (Penstemon strictus) are the only released penstemons noted to have any forage value. All other varieties are considered fair to poor palatability and considered to be only incidental forage value. All species provide diversity to the seeded plant community. Erosion control/reclamation: All species are mentioned for their value in mixes for erosion control and beautification values. Wildlife: Penstemons are considered desirable forages for deer, antelope, and birds either as herbage or seed. They may also provide some cover for selected small bird species. They provide diversity to the plant community.

Status

© Sara Hanna Northern New Mexico Wildflowers Please consult the PLANTS Web site and your State Department of Natural Resources for this plant’s current status, such as, state noxious status and wetland indicator values.

Description

General: Figwort Family (Scrophulariaceae). Penstemon or beardtongue species are perennial forbs or sub-shrubs to shrubs with attractive flowers. They are short to long-lived. Penstemons have opposite, entire, or toothed leaves. They have several stalked flowers or flower clusters that are borne in the axils of the upper leaves or leaflike bracts. The tubular corolla is strongly to distinctly two-lipped at the mouth with a two-lobbed upper lip and a three-lobbed lower lip. There are 4 anther-bearing (fertile) stamens and a single sterile stamen or staminodia that is often hairy at the tip. The fruit is a many-seeded capsule. Penstemon strictus is a perennial herb growing 12 to 36 in tall. It has one to few stems arising from a thick crown. The leaves are entire and narrowly oblanceolate. The flowers are deep blue with parts of the corolla violet and whitish. The staminode is sparsely bearded to glabrous. This species likes full sun to partial shade, well drained soils, and blooms in the summer. A good line drawing and identification key is available in Cronquist et al. (1984).

Distribution

This species is found in the southern Rockies. Except for one minor species, the genus Penstemon does not occur naturally outside of North America. For current distribution, please consult the Plant Profile page for this species on the PLANTS Web site.

Establishment

Adaptation: Penstemons do best on well-drained soils. Most ecotypes do well on infertile, disturbed soils. They have excellent cold winter and drought tolerance. They will tolerate weakly saline to weakly acidic sites. They are usually found in open areas, but will tolerate semi-shaded conditions. They are not tolerant of fire, but are fire resistant due to leaves staying green with relatively high moisture content during the fire season. Rocky Mountain penstemon is often associated with sagebrush in pinyon-juniper woodland, oak scrub, and openings of ponderosa pine and spruce-aspen forest from 2200 to 3200 m in elevation (Cronquist et al. 1984). Planting: These species should be seeded with a drill or broadcast at a depth of 1/4 inch or less into a firm seedbed. Ideal seeding depth is 1/8 inch. Penstemons are not recommended for single species seeding. The full seeding rate (not recommended) for these forbs-shrubs is 1.5 to 3 pounds Pure Live Seed (PLS) per acre or 20 to 26 PLS per square foot (varies somewhat by species). Some recommend 4 lbs PLS per acre for Rocky Mountain penstemon. There are an estimated 478,000 seeds per pound for this species. When used as a component of a mix, adjust to percent of mix desired. For mined lands and other harsh critical areas, doubling the seeding rate component of penstemon is not required. The best seeding results are obtained from seeding in very early spring (because of grass component of mix) on heavy to medium textured soils and in late fall on medium to light textured soils. Late summer (August - mid September) seeding is not recommended. Dormant fall seedings (preferred seeding period for penstemons) will pre-chill seed and reduce seed dormancy which is very strong in some species. Mulching, irrigation, and weed control all benefit stand establishment. Seedling vigor is good, but not as good as most grasses. Germination may not occur until the second growing season. Flowering should not be expected until at least the second growing season. Stands may require weed control measures during establishment. Because penstemons are broadleaf, use of 2,4-D is not recommended. Mow weeds at or prior to their bloom stage. Grasshoppers and other insects may also damage new stands and pesticides may be needed.

Management

Growth of penstemons begins in early spring and flowers appear in May through July depending on species. Weed control and removal of very competitive species may improve chance of establishment. Damage from wildlife and rodents may occur and they may need to be controlled. Disease problems are minimal except under irrigation. Under irrigation, fusarium wilt can be a problem. Environmental Concerns: Penstemon species establish and spread slowly via seed distribution. They are not considered "weedy" or invasive species, but can spread into adjoining vegetative communities under ideal climatic and environmental conditions. They coexist with other native species and add biodiversity to those plant communities.

Seed Production

Penstemons should be seeded in at least 36-inch rows at the rate of 2,5 pounds PLS per acre to allow mechanical weed control, The use of weed barrier material may be an alternative to allow closer spacing, They should be seeded in late fall or early winter unless seed is stratified under cool moist conditions, "The Clearwater Selection" of alpine penstemon does not require stratification, Transplants by dividing the base of older plants or from greenhouse starts can also be successfully used to establish seed fields, Seed is generally harvested by hand stripping or by combine, Seed is mature when capsules are dry and seed is hard and dark in color, Flowering is indeterminate with mature capsules and flowers present at harvest period, Multiple harvest periods (by hand) may be necessary to maximize seed collection, Use soil moisture sensors to measure the soil moisture of Penstemon strictus Benth. ssp. angustus Pennell., Some seed will shatter once capsules open, but the capsule is upright and tends to hold seed very well, Seed can be separated from the capsule by use of a hammer mill or barley de-bearder followed by air screening, Cleaned seed should be allowed to dry and then stored in a cool dry area, An after-ripening period of 3 to 4 months is required, Seed retains viability for several years under these conditions, Cultivars, Improved and Selected Materials (and area of origin) Please check the Vendor Database, expected to be on-line through the PLANTS Web site in 2001 by clicking on Plant Materials, Foundation and registered seed is available for each variety through the appropriate state Crop Improvement Association or commercial sources to grow certified seed, 'Bandera' Rocky Mountain penstemon was developed from seed originally collected in Torrance County, New Mexico, It was developed by Los Lunas Plant Materials Center and released by New Mexico AES, New Mexico State Highway Department, Colorado State University, and the PMC in 1973, It is a native, perennial, cool-season forb with a few large underground stems and many fine roots, The leaves are long and often curl or are reflexed, It is 8 to 23 inches tall and flowers are solitary, on one side of the raceme, The flowers are showy blue to purple and occur primarily between mid May and the end of June, It is best adapted to medium to sandy to rocky textured, weakly acidic to alkaline, well-drained soils, 15 to 20 inch rainfall areas, at 6,000 to 10,000 feet elevation, Its intended uses are for erosion control, diversity, and beautification, Certified seed is available and Los Lunas PMC maintains breeder seed, Other penstemons known to be available on the commercial market include the following: Alpine penstemon (P, venustus) is a selected release from seed originally collected in the Clearwater River drainage, Idaho, 'Cedar' (P, palmeri) Palmer penstemon was developed from seed originally collected near Cedar City in Iron County, Utah, Firecracker penstemon (P, eatonii) is a selected release from seed originally collected near Richfield, Utah, A number of other penstemons are seeded primarily for soil stabilization on depleted, disturbed, and erosive areas for erosion control and as ornamentals, These include low penstemon (Penstemon humilis), Rydberg penstemon (Penstemon rydbergii), and thickleaf penstemon (Penstemon pachyphyllus), No cultivars releases are known to have have been made, Please check the PLANTS database for the names of native penstemons in your state, then check with your area native plant nurseries for their availability, Contact your local Natural Resources

Conservation

Service (formerly Soil Conservation Service) office for more information. Look in the phone book under ”United States Government.” The Natural Resources Conservation Service will be listed under the subheading “Department of Agriculture.”

References

Cronquist, A., A.H. Holmgren, N.H. Holmgren, J.L. Reveal, & P.K. Holmgren 1984. Intermountain flora. Volume Four. Subclass Asteridae (except Asteraceae). The New York Botanical Garden, Bronx, New York. Crosswhite, F.S. 1967. Revision of Penstemon section Habroanthus (Scrophulariaceae). I: Conspectus, II: series Speciosi, and III: series Virgati. American Midland Naturalist 77:1.41. Hanna, S. 2000. Northern New Mexico wildflowers. Version: 000427. New Mexico Highlands University, Las Vegas, New Mexico. <http://venus.nmhu.edu/~english/flower/default.htm> Holmgren, N.H. 1971. A new species of Penstemon from Nye Co., Nevada. Aliso 7:351-356. ________, 1978a. An overlooked new species of Penstemon (Scropulariaceae from the Great Basin. Brittonia 30:334-339. ________, 1978b. Three new species of Penstemon (Scrophulariaceae from the Intermountain region. Brittonia 30:416-425. ________, 1979a. Nomenclatural changes in some Intermountain penstemons (Scrophulariaceae). Brittonia 31:104-107. ________, 1979b. New penstemons (Scrophulariaceae) from the Intermountain region. Brittonia 31:217-242. ________, 1979c. Subgeneric and sectional names for Intermountain Penstemon (Scrophulariaceae). Brittonia 31:358-364. ________, 1980. A new Penstemon (Scrophulariaceae) from northwestern Arizona. Brittonia 32:326-329. Keck, D.D. 1932. Studies in Penstemon. A systematic treatment of the section Saccanthera. Univ. Calif. Publication in Botany 16:367-426. ________, 1937a. Studies in Penstemon IV. The section Ericopsis. Bulletin of the Torrey Botanical Club 64:357-381. ________, 1937b. Studies in Penstemon V. The section Peltanthera. American Midland Naturalist 18:790-829. ________, 1938. Studies in Penstemon VI. The section Autator. Bulletin of the Torrey Botanical Club 65:233-255. ________, 1940. Studies in Penstemon VII. The subsections Gairdneriani, Deusti, and Arenarii of the Graciles, and miscellaneous new species. American Midland Naturalist 23:594-616. ________, 1945. Studies in Penstemon-VIII. A cyto-taxonomic account of the section Spermunculus. American Midland Naturalist 33:128-206. Pennell, F.W. 1920. Penstemon. IN: Scrophulariaceae of the central Rocky Mountain states. Contributions U.S. Natl. Herbarium 20:325.381. USDA, NRCS 2000. The PLANTS database. Version: 000502. <http://plants.usda.gov>. National Plant Data Center, Baton Rouge, Louisiana

Plant Traits

Growth Requirements

Cold Stratification RequiredNo
Hedge ToleranceNone
Frost Free Days, Minimum120
Frost Free Days, Minimum120
Fire ToleranceMedium
Fire ToleranceLow
Fertility RequirementMedium
Fertility RequirementLow
Drought ToleranceMedium
Drought ToleranceMedium
Cold Stratification RequiredNo
Temperature, Minimum (°F)-33
CaCO3 ToleranceMedium
CaCO3 ToleranceLow
Anaerobic ToleranceNone
Anaerobic ToleranceNone
Adapted to Medium Textured SoilsYes
Adapted to Medium Textured SoilsNo
Adapted to Fine Textured SoilsNo
Adapted to Fine Textured SoilsNo
Adapted to Coarse Textured SoilsYes
Adapted to Coarse Textured SoilsYes
Moisture UseLow
Temperature, Minimum (°F)-33
Shade ToleranceIntolerant
Shade ToleranceIntermediate
Salinity ToleranceNone
Salinity ToleranceNone
Root Depth, Minimum (inches)14
Root Depth, Minimum (inches)10
Precipitation, Minimum14
Precipitation, Minimum14
Precipitation, Maximum26
Precipitation, Maximum24
Planting Density per Acre, Minim2700
Planting Density per Acre, Maxim19000
pH, Minimum6.0
pH, Minimum6.0
pH, Maximum8.0
pH, Maximum7.5
Moisture UseMedium
Hedge ToleranceNone


Morphology/Physiology

Active Growth PeriodSpring and Summer
ToxicitySlight
ToxicityNone
Shape and OrientationErect
Shape and OrientationErect
Fall ConspicuousYes
Foliage Porosity WinterPorous
Foliage Porosity SummerPorous
Foliage Porosity SummerModerate
Foliage ColorGreen
Foliage ColorGreen
Flower ConspicuousYes
Flower ConspicuousNo
Flower ColorRed
Flower ColorPurple
Fire ResistantNo
Fire ResistantNo
Resprout AbilityNo
Fall ConspicuousNo
Coppice PotentialNo
Coppice PotentialNo
C:N RatioMedium
C:N RatioMedium
BloatNone
BloatNone
After Harvest Regrowth RateSlow
After Harvest Regrowth RateSlow
Active Growth PeriodSpring and Summer
Resprout AbilityNo
Foliage TextureCoarse
Nitrogen FixationNone
Nitrogen FixationNone
Low Growing GrassNo
Low Growing GrassNo
LifespanModerate
LifespanLong
Leaf RetentionNo
Leaf RetentionNo
Known AllelopathNo
Known AllelopathNo
Height, Mature (feet)2.5
Growth RateRapid
Height, Mature (feet)2.0
Foliage Porosity WinterPorous
Foliage TextureMedium
Fruit/Seed ColorBrown
Fruit/Seed ColorBrown
Fruit/Seed ConspicuousNo
Fruit/Seed ConspicuousYes
Growth FormBunch
Growth FormStoloniferous
Growth RateModerate


Reproduction

Propagated by CuttingsNo
Propagated by SeedYes
Propagated by SeedYes
Propagated by SodNo
Propagated by SodNo
Propagated by SprigsNo
Propagated by SprigsNo
Propagated by TubersNo
Propagated by TubersNo
Seed per Pound286000
Fruit/Seed Period EndSummer
Seed per Pound489888
Seed Spread RateModerate
Seed Spread RateSlow
Seedling VigorHigh
Seedling VigorLow
Small GrainNo
Small GrainNo
Vegetative Spread RateModerate
Vegetative Spread RateNone
Propagated by CormNo
Propagated by CuttingsNo
Bloom PeriodLate Spring
Bloom PeriodLate Spring
Commercial AvailabilityRoutinely Available
Commercial AvailabilityRoutinely Available
Fruit/Seed AbundanceMedium
Fruit/Seed AbundanceMedium
Fruit/Seed Period BeginSpring
Fruit/Seed Period BeginSpring
Fruit/Seed Period EndSummer
Fruit/Seed PersistenceNo
Propagated by Bare RootNo
Propagated by Bare RootNo
Propagated by BulbNo
Propagated by BulbNo
Propagated by ContainerNo
Propagated by ContainerYes
Propagated by CormNo
Fruit/Seed PersistenceNo


Suitability/Use

Palatable Browse AnimalMedium
Palatable Graze AnimalLow
Palatable Graze AnimalMedium
Palatable HumanNo
Palatable HumanNo
Post ProductNo
Post ProductNo
Protein PotentialLow
Protein PotentialMedium
Pulpwood ProductNo
Pulpwood ProductNo
Veneer ProductNo
Veneer ProductNo
Palatable Browse AnimalMedium
Nursery Stock ProductYes
Nursery Stock ProductNo
Naval Store ProductNo
Naval Store ProductNo
Lumber ProductNo
Lumber ProductNo
Fodder ProductYes
Fodder ProductNo
Christmas Tree ProductNo
Christmas Tree ProductNo
Berry/Nut/Seed ProductNo
Berry/Nut/Seed ProductNo

<i>Penstemon strictus</i> Benth. ssp. angustus Pennell

<i>Penstemon strictus</i> Benth. ssp. angustus Pennell

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