Bush Muhly

Scientific Name: Muhlenbergia porteri Scribn. ex Beal

Classification: Plantae/ Tracheobionta / Spermatophyta / Magnoliophyta / Liliopsida / Commelinidae / Cyperales / Poaceae / Gramineae / Muhlenbergia Schreb./ Muhlenbergia porteri Scribn. ex Beal

Bush Muhly
General Information
Usda SymbolMUPO2
GroupMonocot
Life CyclePerennial
Growth HabitsGraminoid
Native LocationsMUPO2

Plant Guide

Description

General: Bush muhly is a native perennial, warm-season, shrubby bunchgrass that grows 1.5 to 3 ft tall and wide. The common name refers to the bush-like appearance of plants; the plants are also often seen spreading up through the branches of bushes. Bush muhly has a semi-woody, decumbent base that is not rhizomatous. Culms are wiry, bent at the nodes, branching and intertwined. Leaf blades are very narrow, up to 3 inches long, and may be flat or folded. The inflorescence is an open panicle 2 to 4 inches long with thread-like branches. The spikelets may have a purple color in cool weather. Bush muhly begins growth in late winter or early spring, and flowers from early spring through summer (Kearney and Peebles, 1964; Hitchcock, 1971; Felger et al., 2014; Peterson, 2012; USU, 2017). Distribution: Bush muhly occurs in arid regions of California, Nevada, Utah, Colorado, Arizona, New Mexico, Oklahoma, Texas, and in Northern Mexico. For current distribution, please consult the Plant Profile page for this species on the PLANTS Web site. Habitat: Bush muhly grows on mesas, rocky slopes, and along drainages. It typically occurs at elevations between 2000 and 6000 feet above sea level. Bush muhly is most commonly found in areas that are protected from grazing (within shrubs, between boulders). It was formerly a dominant grass in southern Arizona but is now generally found in limited quantities throughout its range due to overgrazing (Ruyle and Young, 2003).

Adaptation

Bush muhly is a C4 grass adapted to hot, arid conditions. It is considered a climax species (Glendening et al., 1955). Bush muhly is tolerant of a wide variety of soil textures. It tolerates calcium carbonate but not soil salinity. During dry and cool periods bush muhly may behave like a shrub, with its leaves dying back to the nodes (Van Devender and Dimmitt, 2000). Under favorable conditions, leaves and inflorescences can quickly regrow from the nodes. Fire and hard frost cause the plants to die back to the crown, regrowing when conditions improve (Van Devender and Dimmitt, 2000). Bush muhly has a higher water use efficiency than creosote bush (Castellanos-Pérez et al, 2008) or mesquite seedlings (Glendening et al., 1955), giving it the ability to inhibit the growth of these competitors. Bush muhly can survive under mesquite canopies (Jacoby et al., 1982). Associated plants include velvet mesquite (Prosopis velutina), ironwood (Olneya tesota), catclaw acacia (Senegalia greggii), creosote bush (Larrea tridentata), saguaro (Carnegiea gigantea), Joshua tree (Yucca brevifolia), banana yucca (Yucca baccata), desert spoon (Dasylirion wheeleri), fourwing saltbush (Atriplex canescens), sagebrush (Artemisia tridentata), brittlebush (Encelia sp.), broom snakeweed (Gutierrezia sarothrae), tobosagrass (Pleuraphis mutica), purple threeawn (Aristida purpurea), sand dropseed (Sporobolus cryptandrus), and cane bluestem (Bothriochloa barbinodis).

Uses

Grazing: Bush muhly is highly palatable remaining green through most of the year when moisture is available. However, its branching habit makes it susceptible to damage by overgrazing (Utah State University, 2017). Formerly abundant, it is now more commonly found in places where livestock cannot reach it. Bush muhly was grazed out of large areas of Organ Pipe, AZ, prior to the creation of the National Monument. After cattle were removed, it increased to become common within the Bush muhly. (Photo by Heather Dial, USDA-NRCS Tucson Plant Materials Center) Plant Guide Monument (Felger et al., 2014). Wildlife: Desert tortoises eat bush muhly and may also use the plants for shelter (Felger et al., 2014; McArthur et al. 1994). Cactus wrens commonly gather bush muhly for nesting material, probably dispersing the seeds in the process (Milton et al 1999).

Ethnobotany

The Tohono O’odham word for bush muhly, ku:kpadag, means “plant formerly used to close water containers” (Hill and Zepeda, 1994), suggesting that clumps of bush muhly were used as stoppers.

Status

Threatened or Endangered: No. Wetland Indicator: Bush muhly is an upland (UPL) plant. Weedy or Invasive: Bush muhly is not known to be weedy or invasive. Please consult the PLANTS Web site (http://plants.usda.gov/) and your state’s Department of Natural Resources for this plant’s current status (e.g., threatened or endangered species, state noxious status, and wetland indicator values).

Planting Guidelines

Bush muhly typically reproduces by seed. The recommended seeding rate for bush muhly is 0.45 pure live seed (PLS) pounds per acre if planted with a drill and approximately 0.9 PLS pounds per acre if seed is broadcast. There are approximately 2,424,000 seeds of bush muhly in a pound (USDA-PLANTS Database). For containerized production, no pre-treatment methods are necessary as seeds are non-dormant (Baskin and Baskin, 2002).

Management

Protect bush muhly from overgrazing. When possible, defer grazing in late summer every second or third year allowing the plants to set seed (Ruyle and Young, 2003; Allison and Ashcroft, 2011).

Pests and Potential Problems

None known, Use soil moisture sensors to measure the soil moisture of Bush Muhly.,

Environmental Concerns

Concerns

Concerns

None known.

Seed and Plant Production

Plant Production

Plant Production

Bush muhly seed should be planted in the early spring into a firm, weed free seedbed at a depth of 1/4 inch. New seedings should be irrigated to maintain a moist soil surface and to avoid soil crusting. Plugs should be planted with 40-48 inches between rows, and 36-40 inch within-row spacing. Control weeds as needed with an appropriate herbicide after plants have developed at least 3-5 leaves. Apply nutrients according to soil test results. Irrigate established fields approximately every four weeks during the growing season. Irrigated fields produce seed in summer. Harvest with a seed stripper or combine. Clean harvested seed by processing with a hammermill or brush machine to break up the panicles; follow with air screening. Cultivars, Improved, and Selected Materials Limited amounts of this species are available on the commercial market. Currently, none are listed as cultivars, improved or selected materials.

Literature Cited

Allison, C.D. and N. Ashcroft. 2011. New Mexico Range Plants. Circular 374. [Online]. Available at https://aces.nmsu.edu/pubs/_circulars/CR374/ (accessed on 30 May 2019). College of Agricultural, Consumer and Environmental Sciences, New Mexico State University, Las Cruces, New Mexico. Baskin, J.M. and C.C. Baskin. 2002. Propagation protocol for production of container (plug) Muhlenbergia porteri Scribn. plants [Online]. Available at http://NativePlantNetwork.org (accessed on 13 March 2019). In: Native Plant Network. USDA, Forest Service, National Center for Reforestation, Nurseries, and Genetic Resources. University of Kentucky, Lexington, Kentucky. Bush muhly production fields at the USDA-NRCS Tucson Plant Materials Center. (Photo by Heather Dial)

http://plants.usda.gov/ Felger, R.S., S. Rutman and J. Malusa. 2014. Ajo Peak to Tinajas Altas: A flora of southwestern Arizona. Part 6. Poaceae – grass family. Phytoneuron 2014-35: 1-139. Hill, J.H. and O. Zepeda. 1994. Tohono O’odham (Papago) plurals. In Estudios de Lingüística y Sociolingüística. G.López Cruz and J.L. Moctezuma Zamarrón, eds. Departamento de Letras y Lingüística, División de Humanidades y Bellas Artes, Universidad De Sonora, Instituto Nacional de Antropología e Historia, Hermosillo, Sonora, Mexico. Hitchcock, A.S. 1971. Manual of the Grasses of the United States (Second Edition, Rev. by Agnes Chase). Dover Publications, Inc., New York, NY. Jacoby, P.W., C.H. Meadors, M.A. Foster, and F.S. Hartmann. 1982. Honey mesquite control and forage response in Crane County, Texas. Journal of Range Management. 35:424-426. Kearney, T.H. and R.H. Peebles. 1964. Arizona Flora with Supplement. University of California Press, Berkeley and Los Angeles, CA. McArthur, E.D., S.C. Anderson and B.L. Webb. 1994. Nutritive quality and mineral content of potential desert tortoise food plants. Research Paper INT-473, Intermountain Research Station. U.S. Forest Service, United States Department of Agriculture. Milton, S. J., W. J. Dean, G. H. Kerley, M. T. Hoffman, and W G. Whitford. 1999. Dispersal of seeds as nest material by the cactus wren. Southwestern Naturalist 43(4):449-452. Peterson, P.M. 2012. Muhlenbergia porteri, in Jepson Flora Project (eds.). Jepson eFlora [Online] Available at http://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=34110 (accessed on March 13, 2019). Ruyle, G.B. and D.J. Young, eds. 2003. Common Arizona range grasses: Their description, forage value and management. Cooperative Extension, College of Agriculture and Life Sciences. The University of Arizona, Tucson, Arizona. USDA-NRCS. 2019. The PLANTS Database [Online]. Available at http://plants.usda.gov (accessed on March 13, 2019). National Plant Data Team, Greensboro, NC. Utah State University Extension. 2017. Bush muhly. Range Plants of Utah [Online]. Available at https://extension.usu.edu/rangeplants/grasses-and-grasslikes/bush-muhly (accessed on January 10, 2019). Van Devender, T.R. and M.A. Dimmitt. 2000. Desert Grasses. In A Natural History of the Sonoran Desert, 1st ed. (eds.) S.J. Phillips and P.W. Comus. Arizona-Sonora Desert Museum, Tucson, AZ. Citation Wolf, M. 2019. Plant Guide for bush muhly (Muhlenbergia porteri). USDA-Natural Resources Conservation Service, Tucson Plant Materials Center. Tucson, AZ 85705. Published June 2019 For more information about this and other plants, please contact your local NRCS field office or Conservation District at http://www.nrcs.usda.gov/ and visit the PLANTS Web site at http://plants.usda.gov/ or the Plant Materials Program web site: http://plant-materials.nrcs.usda.gov. PLANTS is not responsible for the content or availability of other Web sites.
http://www.nrcs.usda.gov/
http://plants.usda.gov/
http://plant-materials.nrcs.usda.gov/ In accordance with Federal civil rights law and U.S. Department of Agriculture (USDA) civil rights regulations and policies, the USDA, its Agencies, offices, and employees, and institutions participating in or administering USDA programs are prohibited from discriminating based on race, color, national origin, religion, sex, gender identity (including gender expression), sexual orientation, disability, age, marital status, family/parental status, income derived from a public assistance program, political beliefs, or reprisal or retaliation for prior civil rights activity, in any program or activity conducted or funded by USDA (not all bases apply to all programs). Remedies and complaint filing deadlines vary by program or incident. Persons with disabilities who require alternative means of communication for program information (e.g., Braille, large print, audiotape, American Sign Language, etc.) should contact the responsible Agency or USDA's TARGET Center at (202) 720-2600 (voice and TTY) or contact USDA through the Federal Relay Service at (800) 877-8339. Additionally, program information may be made available in languages other than English. To file a program discrimination complaint, complete the USDA Program Discrimination Complaint Form, AD-3027, found online at How to File a Program Discrimination Complaint and at any USDA office or write a letter addressed to USDA and provide in the letter all of the information requested in the form. To request a copy of the complaint form, call (866) 632-9992. Submit your completed form or letter to USDA by: (1) mail: U.S. Department of Agriculture, Office of the Assistant Secretary for Civil Rights, 1400 Independence Avenue, SW, Washington, D.C. 20250-9410; (2) fax: (202) 690-7442; or (3) email: program.intake@usda.gov. USDA is an equal opportunity provider, employer, and lender.

Plant Traits

Growth Requirements

Temperature, Minimum (°F)12
Adapted to Coarse Textured SoilsYes
Adapted to Fine Textured SoilsYes
Adapted to Medium Textured SoilsYes
Anaerobic ToleranceNone
CaCO3 ToleranceNone
Cold Stratification RequiredNo
Drought ToleranceHigh
Fertility RequirementMedium
Fire ToleranceHigh
Frost Free Days, Minimum220
Hedge ToleranceNone
Moisture UseMedium
pH, Maximum8.0
pH, Minimum6.2
Precipitation, Maximum12
Precipitation, Minimum5
Root Depth, Minimum (inches)12
Salinity ToleranceLow
Shade ToleranceIntermediate


Morphology/Physiology

After Harvest Regrowth RateSlow
ToxicityNone
Shape and OrientationDecumbent
Nitrogen FixationNone
Resprout AbilityNo
Active Growth PeriodSpring, Summer, Fall
BloatNone
C:N RatioMedium
Coppice PotentialNo
Fall ConspicuousNo
Fire ResistantNo
Flower ColorYellow
Flower ConspicuousNo
Foliage ColorGreen
Foliage Porosity SummerDense
Foliage TextureFine
Low Growing GrassNo
LifespanLong
Leaf RetentionYes
Known AllelopathNo
Height, Mature (feet)2.8
Growth RateModerate
Growth FormSingle Crown
Fruit/Seed ConspicuousNo
Fruit/Seed ColorBrown
Foliage Porosity WinterModerate


Reproduction

Vegetative Spread RateSlow
Small GrainNo
Seedling VigorMedium
Seed Spread RateSlow
Seed per Pound2424000
Fruit/Seed PersistenceNo
Propagated by TubersNo
Propagated by SprigsNo
Propagated by SodNo
Propagated by SeedYes
Propagated by CormNo
Propagated by ContainerNo
Propagated by BulbNo
Propagated by Bare RootNo
Fruit/Seed Period EndSummer
Fruit/Seed Period BeginSummer
Fruit/Seed AbundanceLow
Commercial AvailabilityContracting Only
Bloom PeriodLate Spring
Propagated by CuttingsNo


Suitability/Use

Veneer ProductNo
Pulpwood ProductNo
Protein PotentialMedium
Post ProductNo
Palatable HumanNo
Palatable Graze AnimalHigh
Palatable Browse AnimalHigh
Nursery Stock ProductNo
Naval Store ProductNo
Lumber ProductNo
Fodder ProductYes
Christmas Tree ProductNo
Berry/Nut/Seed ProductNo

Bush Muhly

Bush Muhly

Bush Muhly

Bush Muhly